cOVER

Contents

Preface

Acknowledgements

Introduction

What makes a beetle?

Anatomy

Reproduction and development

Food and survival

Higher taxonomy

Family descriptions

Cupedidae Reticulated beetles

Ommatidae Ommatid beetles

Sphaeriusidae Minute bog beetles

Gyrinidae Whirligig beetles

Haliplidae Crawling water beetles

Noteridae Burrowing water beetles

Hygrobiidae Screech beetles

Dytiscidae Predaceous diving beetles

Rhysodidae Wrinkled bark beetles

Carabidae Ground beetles

Hydrophilidae Water scavenger beetles

Histeridae Hister beetles

Hydraenidae Minute moss beetles

Ptiliidae Feather-winged beetles

Leiodidae Small fungus beetles

Scydmaenidae Minute ant-like beetles

Silphidae Carrion or burial beetles

Staphylinidae Rove beetles

Lucanidae Stag beetles

Passalidae Passalid or bess beetles

Trogidae Hide beetles

Bolboceratidae Bolboceratid beetles

Hybosoridae Scavenger scarab beetles

Scarabaeidae Scarab beetles

Aclopinae Aclopine scarab beetles

Aphodiinae Aphodiine dung beetles

Scarabaeinae Dung beetles

Dynastinae Rhinoceros or elephant beetles

Melolonthinae Chafers

Rutelinae Shining leaf chafers

Cetoniinae Flower chafers

Scirtidae Marsh beetles

Clambidae Minute spherical beetles

Dascillidae Soft-bodied plant beetles

Rhipiceridae Feather- or fan-horned beetles

Buprestidae Jewel or metallic wood-boring beetles

Byrrhidae Pill beetles

Callirhipidae Callirhipid beetles

Chelonariidae Chelonarid beetles

Elmidae Riffle beetles

Heteroceridae Variegated mud-loving beetles

Limnichidae Minute marsh-loving beetles

Psephenidae Water-penny beetles

Ptilodactylidae Toe-winged beetles

Brachypsectridae Texas beetles

Eucnemidae False click beetles

Throscidae Throscid beetles

Elateridae Click beetles

Lycidae Net-winged beetles

Lampyridae Fireflies or flashing fireflies

Rhinorhipidae Rhinorhipid beetles

Cantharidae Soldier beetles

Nosodendridae Wounded-tree beetles

Dermestidae Dermestid beetles

Bostrichidae Auger or shot-hole borers

Anobiidae Furniture and spider beetles

Lymexylidae Ship-timber beetles

Trogossitidae Bark-gnawing beetles

Cleridae Checkered beetles

Phycosecidae Phycosecid beetles

Melyridae Soft-winged flower beetles

Nitidulidae Sap beetles

Silvanidae Silvanid flat bark beetles

Passandridae Parasitic flat bark beetles

Cucujidae Flat bark beetles

Laemophloeidae Lined flat bark beetles

Phalacridae Shining flower or shining mould beetles

Cryptophagidae Silken fungus beetles

Erotylidae Pleasing fungus beetles

Languriinae Lizard beetles

Bothrideridae Dry bark beetles

Endomychidae Handsome fungus beetles

Discolomatidae Tropical log beetles

Coccinellidae Lady or ladybird beetles

Corylophidae Minute fungus beetles

Latridiidae Minute brown scavenger beetles

Mycetophagidae Hairy fungus beet

Ciidae Minute tree-fungus beetles

Melandryidae False darkling beetles

Mordellidae Pin-tail or tumbling flower beetles

Rhipiphoridae Wedge-shaped beetles

Zopheridae

Zopherinae Ironclad beetles

Colydiinae Colydiine beetles

Ulodidae Ulodid beetles

Tenebrionidae Darkling beetles

Lagriinae Long-jointed bark beetles

Tenebrioninae True darkling beetles

Alleculinae Comb-clawed beetles

Diaperinae Diaperine beetles

Stenochiinae Stenochiine beetles

Oedemeridae False blister beetles

Meloidae Blister or oil beetles

Mycteridae Palm and flower beetles

Salpingidae Narrow-waisted bark beetles

Anthicidae Ant-like flower beetles

Aderidae Ant-like leaf beetles

Scraptiidae False flower beetles

Cerambycidae Longhorn or longicorn beetles

Chrysomelidae Leaf beetles

Bruchinae Seed weevils

Cassidinae

(Cassidoid group) Tortoise beetles

(Hispoid group) Spiny leaf beetles

Chrysomelinae Chrysomelin leaf beetles

Criocerinae Shining leaf beetles

Cryptocephalinae Cylindrical leaf beetles

Eumolpinae Oval leaf or leaf monkey beetles

Galerucinae Galerucine and flea leaf beetles

Sagrinae Kangaroo beetles

Nemonychidae Pine flower weevils

Anthribidae Fungus weevils

Belidae Belid weevils

Attelabidae Leaf-rolling weevils

Brentidae Straight-snouted weevils

Curculionidae Weevils or snout beetles

Endnotes

Glossary

Index of common names

Index of scientific names